at the Department of Zoology. Closing date: 1 March 2018.

The Department of Zoology is a vibrant international community, consisting of five interactive and collaborative divisions: Ecology, Ethology, Functional Morphology, Population Genetics, and Systematics and Evolution. The advertised PhD will be part of the Division of Ecology.

Project description
A PhD position focused on sexual selection and sexual conflict is available in Rhonda Snook’s group, recently relocated to the Department of Zoology, Stockholm University. Sexual selection and sexual conflict results in the rapid (co)evolution of male and female reproductive traits, such as female remating behavior and male ejaculate proteins, which may be critical in local adaptation and speciation. However, identifying the ‘locus of sexual selection’ (i.e., the sequence variants) that cause differences in sexually selected traits within and between populations, and between the sexes sharing a genome, is rudimentary, hampering the understanding of the role of sexual selection in trait elaboration, diversification and speciation. The PhD project will address this knowledge gap by targeting an easily quantifiable female behavioral trait (sperm ejection) and male ejaculate traits known to undergo rapid evolution (seminal fluid proteins), both of which have profound effects on reproductive success. The research will uncover the genetic architecture of the coevolution between sperm ejection and SFPs using sequenced lines of Drosophila melanogaster and experimentally evolved populations of D. pseudoobscura in which sexual selection has been manipulated. Phenotypic assays, forward and reverse molecular genetic techniques, and proteomics and downstream bioinformatic analyses will be used.

Qualification requirements
In order to meet the general entry requirements, the applicant must have completed a second-cycle degree, completed courses equivalent to at least 240 higher education credits, of which 60 credits must be in the second cycle, or have otherwise acquired equivalent knowledge in Sweden or elsewhere.

In order to meet the specific entry requirements, the general syllabus for doctoral studies in the field of Ecology stipulates that applicants must have completed a research degree (e.g. Master’s), or have passed at least 120 hp (2 years) of biological studies, including an approved independent project of at least 30 hp at advanced level (“examensarbete”) within evolutionary ecology, molecular genetics/bioinformatics, or a similar subject. Applicants who have in principle acquired the corresponding competence in Sweden, or abroad, are also qualified.

The qualification requirements must be met by the deadline for applications.

The selection among the eligible candidates will be based on their capacity to benefit from the training. I am looking for candidates with a strong interest in evolutionary ecology, ideally with knowledge of molecular genetics/bioinformatics. Prior training in insect culture would be helpful but not essential. Applicants should be well-organized, good communicators, and happy to work in a team as well as independently. The work will be lab-oriented. The skills required for the project will include large scale experiments for collection of phenotypic data, large scale insect culture, molecular genetics techniques, and subsequent analysis of complex phenotypic and genomic data. The project will require hard work in the lab but in a convivial and interactive atmosphere, allowing for additional collaborations within the group, the department, and adjacent biology departments (DEEP, MBW) and the Bolin Centre for Climate Research. The ability to shape the project will provide excellent opportunities for personal initiatives and development towards a successful academic career. The working language of the group is English.

Admission Regulations for Doctoral Studies at Stockholm University are available at:

Terms of employment
Only a person who will be or has already been admitted to a third-cycle programme may be appointed to a doctoral studentship.

The term of the initial contract may not exceed one year. The employment may be extended for a maximum of two years at a time. However, the total period of employment may not exceed the equivalent of four years of full-time study.

Doctoral students should primarily devote themselves to their own education, but may engage in teaching, research, and administration corresponding to a maximum of 20 % of a full-time position.

Please note that admission decisions cannot be appealed.

Stockholm University strives to be a workplace free from discrimination and with equal opportunities for all.

For more information, please contact Rhonda Snook,

Union representatives
Ingrid Lander (Saco-S) and Lisbeth Häggberg (Fackförbundet ST and Lärarförbundet), telephone: +46 8 16 20 00 (operator), (SEKO), and PhD student representative,

Apply for the PhD student position at Stockholm University's recruitment system by clicking the "Apply" button. It is the responsibility of the applicant to ensure that the application is complete in accordance with the instructions in the job advertisement, and that it is submitted before the deadline.

Please include the following information with your application

  • Your contact details and personal data
  • Your highest degree
  • Your language skills
  • Contact details for 2–3 references

and, in addition, please include the following documents

  • Cover letter
  • CV – degrees and other completed courses, work experience and a list of degree projects/theses
  • Degree certificates and grades confirming that you meet the general and specific entry requirements (no more than 6 files)
  • Degree projects/theses (no more than 6 files).

The instructions for applicants are available at: Instructions – Applicants.

You are welcome to apply!

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Closing date: 01/03/2018